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Monoterpene alcohols can be further metabolized by yeasts, through esterification or hydrolysis, particularly acetylation of monoterpenes by AATase enzymes (King and Dickinson 2000) or by cleavage of monoterpene esters by esterases or lipases (Lam, Foster and Deinzer 1986

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Therefore, the ability of metformin to enhance GSH levels may be a key mechanism by which it attenuates intestinal ferroptosis