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Since mitochondrial fragmentation is mainly caused by mitochondrial fission and fusion imbalance or mitochondrial apoptosis 29 , we further verified the protein and mRNA expression levels of mitochondrial fission (Drp1), fusion (Opa1 and Mfn2), apoptosis (Tom20 and COX IV) and mitochondrial respiration (SDHB, NDUFV2 and ATP5A1) markers by western blotting and PCR