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This article was submitted to Predictive Toxicology, a section of the journal Frontiers in Pharmacology Disclaimer All claims expressed in this article are solely those of the authors and do not necessarily represent those of their affiliated organizations, or those of the publisher, the editors and the reviewers

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Original and newly shifted R S boundaries are indicated with dashed lines

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Berberine: The Centerpiece Ingredient Berberine is a bioactive alkaloid compound found in several plants, including goldenseal (Hydrastis canadensis), barberry (Berberis vulgaris), and Oregon grape (Mahonia aquifolium)